Degree :
University of Minnesota, Minneapolis, Minnesota
B.S. in Biochemistry (1981)
Undergraduate Research Advisor: Dr. Samuel Kirkwood
University of Minnesota Medical School, Minneapolis, Minnesota
M.D. (1985)
Residency:
Mayo Graduate School of Medicine, Rochester, Minnesota
Medicine Resident (1985 - 1988)
Fellowship:
Duke University Medical School, Durham, North Carolina
Fellow, Division of Allergy, Critical Care and Respiratory Medicine (1989 - 1992)
Fellowship Research Advisors: Dr. James Bonner and Dr. Arnold Brody
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